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Wireless sensor networks (WSNs) have been a widely researched field since the beginning of the 21st century. The field is already maturing, and TinyOS has established itself as the de facto standard WSN Operating System (OS). However, the WSN researcher community is still active in building more flexible, efficient and user-friendly WSN operating systems. Often, WSN OS design is based either on practical requirements of a particular research project or research group’s needs or on theoretical assumptions spread in the WSN community. The goal of this paper is to propose WSN OS design rules that are based on a thorough survey of 40 WSN deployments. The survey unveils trends of WSN applications and provides empirical substantiation to support widely usable and flexible WSN operating system design.