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A necessary ingredient of an ensemble Kalman filter (EnKF) is covariance inflation, used to control filter divergence and compensate for model error. There is an on-going search for inflation tunings that can be learned adaptively. Early in the development of Kalman filtering, Mehra (1970, 1972) enabled adaptivity in the context of linear dynamics with white noise model errors by showing how to estimate the model error and observation covariances. We propose an adaptive scheme, based on lifting Mehra's idea to the non-linear case, that recovers the model error and observation noise covariances in simple cases, and in more complicated cases, results in a natural additive inflation that improves state estimation. It can be incorporated into non-linear filters such as the extended Kalman filter (EKF), the EnKF and their localised versions. We test the adaptive EnKF on a 40-dimensional Lorenz96 model and show the …