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Most applications of wireless sensor networks require reliable and timely data communication with maximum possible network lifetime under low traffic regime. These requirements are very critical especially for the stability of wireless sensor and actuator networks. Designing a protocol that satisfies these requirements in a network consisting of sensor nodes with traffic pattern and location varying over time and space is a challenging task. We propose an adaptive optimal duty-cycle algorithm running on top of the IEEE 802.15.4 medium access control to minimize power consumption while meeting the reliability and delay requirements. Such a problem is complicated because simple and accurate models of the effects of the duty cycle on reliability, delay, and power consumption are not available. Moreover, the scarce computational resources of the devices and the lack of prior information about the topology make it …