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The aim of this paper is to investigate how research partnerships between firms and universities emerge and evolve over time, focusing on the relationship between initial conditions and development trajectories. Based on literature on alliances and a comparative case study, the paper presents propositions to explore the links between initial conditions, modes of collaboration and development pathways. The empirical study indicates that there is no straightforward link between initial conditions and development of partnerships over time. Being embedded in a network makes it easier to form relationships and has a positive influence of the early, set-up phase of collaboration, but has limited impact on long term development of partnerships. The data also illustrates the weak side of networks; they are efficient mechanism for forming relationships but in some cases represent weak commitment to partnerships …