and secondary forests of Puerto Rico. Tree plantations included pine (Pinus caribaea
Morelet) and mahogany (Swietenia macrophylla King) established in the 1930s; 1960s; and
1970s; secondary forests were naturally regenerated in areas adjacent to these plantations.
We found that (1) earthworm density and fresh weight in the secondary forests were twice
those in either of the tree plantations, and did not differ between the plantations, and (2) the …