soil organisms (Woomer et al., 1994; Tan, 2010). The energy requirements of micro and
macro organisms, other than autotrophs and chemotrophs, present in the soil are largely met
by the organic matter added and the native soil organic matter (Monisa and Tahir, 2018).
Soil enzyme activity estimates are often used as indices of microbial activity and soil fertility
(Vaughan and Malcolm, 1985; Gianfreda and Bollag, 1996; Ranjith et al.,