Exceptionally preserved lycopsid remains from an ex situ chert–carbonate cobble found on
the Yorkshire coast, United Kingdom, are here reconstructed as Flemingites arcuatus sp.
nov. Megasporophylls are C shaped and emerge from the cone axis at a relatively acute
angle. Megasporangia are adaxial on sporophylls and typically contain four megaspore
tetrads. Megaspores are described as Lagenicula wellmanii sp. nov.; they are 1.0–1.7 mm in
diameter and have a pronounced gula and numerous vermiform spines with bulbous apices …

On the Cover: Inset, a young sporophyte of Selaginella apoda, a native of moist and shady
areas throughout the United States and Mexico (photograph by Christian Schulz and Ciaran
Moloney). Selaginella apoda presents many advantages as a model species (see Schulz et
al., pp. 693–712, in this issue). Background, at shedding, the bright yellow megaspores of
Selaginella apoda are~ 300 μm in size (photograph by Christian Schulz).