… results support the concept that the cuttlefish patterning responses depend on substrate
spatial scale. We discuss the scale-dependency in the context of cuttlefish camouflage tactics. …

… -scale light and dark patches, again with minimal contrast. Mottle patterns have small or
large-scale … Both uniform and mottle patterns achieve camouflage by the principle of general …

… are key visual cues that guide cephalopod camouflage patterning. Mottle and disruptive …
For readability, we use a scale in which energy is normalized by the maximum value of the …

… patterns are strongly influenced by visual edges in the substrate. The aim of the present study
was to examine how cuttlefish body patterning … The Y-axis is scaled based on the standard …

… We studied here the 2D features of camouflage and therefore refer to them as skin patterns
… We first standardized features by removing mean and scaling to unit variance. We obtained …

… light and dark objects (or with some repetition of small-scale … cues to elicit mottle camouflage
patterns in cuttlefish. Lowering … scale of mottle patterns whereas A2 showed medium scale …

… suggest that body patterning of cuttlefish is scale dependent. … the scale dependency of the
cuttlefish patterning responses … patterning responses depend on substrate spatial scale (Fig. …

… various ways that colours and patterns could be used to improve camouflage. What are now
… as a sample of the background at the spatial scale of the animal, has a long history [2] and …

… into camouflage design. Edge detection is an important aspect of vision, and here we compare
the body patterns … The prints were calibrated for linearity with a grey scale over a range of …

… Moreover, it will also allow us to design effective camouflage patterns for … camouflage when
they were at the same orientation as the stripes in their background at a similar spatial scale. …